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One canonical way for -KG to regulate gene transcription is to act as an epigenetic cofactor for Fe 2+ -dependent and 2-oxoglutarate-dependent dioxygenases (2OGDDs), including TET DNA demethylases and Jumonji C domain-containing histone demethylases (JMJD3 and UTX) 16,42,43

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Based on the evidence that heat-shock proteins (Hsps) can exert neuroprotective effects against oxidative stress-related injury and that nutritional antioxidants are able to upregulate Hsps in neurons, use of nutritional antioxidants such as carnitine/acetyl-L-carnitine has been advocated to counteract the oxidative stress-induced brain damage in Alzheimer's disease [67, 119]

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